After her solitary punishment, Luce questions her memory of what happened to Cam. She tries to retrieve his medical records, but Daniel refuses to help. Meanwhile, Cassie, the only one to escape, encounters a nightmarish creature.

Luce suspects Howson is controlling their memories and steals photos of her and others undergoing medical procedures. While on a hike, Howson learns of Cassie's attack and brings her to Sword and Cross.

Howson captures the creature that attacked Luce in the lake and interrogates it. While Luce comforts Daniel over Cassie's death, Penn uncovers troubling truths about her past. With her father's arrival, Luce and Daniel get the medical files.

Luce learns Daniel hasn't aged in over twenty years. Determined to expose the truth, she plans a breakout with Daniel as proof. However, Howson is tipped off, and the Outcast injures Daniel. Luce finds the body of another Lucinda Price.

Miss Miriam hunts the Outcast while Howson fights to save Daniel. Luce and the others try to escape, but the creature pursues them. Luce learns who betrayed them to Howson and why. Meanwhile, Daniel fades as he recalls a past life with another Luce.

Miss Miriam reveals some secrets of Sword & Cross. Luce faces her own struggles, fearing she'll die if they stay together. Luce has a vision of Penn's death and tries to prevent her from leaving, but someone is waiting for Penn with deadly intent.

Luce believes she was murdered. Daniel confronts Miss Miriam, and Cam faces Howson. Luce seeks answers from Miss Sophia. Daniel saves her, and they reconcile. Luce faces a violent end that could change humanity's future.